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Invasion Land-Air
Apical Meristems & Branching

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Apical Cells and their most recent derivatives constitute the Apical Meristems of Ancestral Plants.

This includes most species in the Hepatophyta, Bryophyta, Psilophyta, Sphenophyta, Lycophyta and Pterophyta.

 

Members of the Coniferophyta and Magnoliophyta have Multicellular Apical Meristems.

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Equisetum Shoot apex with a prominent Apical Cell

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Multicellular Shoot apical Meristem from a Flowering Plant (Magnoliophyta)

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Multicellular Root Apical Meristem from a Gymnosperm

Branching

Ancestral Plants had Isotomous Branching in their Stems and Roots. This produces branches of equal length at each branch point.This is a form of Dichotomous Branching which occurs at the Shoot or Root Apical Meristem.

More Advanced plants had Anisotomous Branching. This produces a short branch and a long branch at each dichotomy. These alternate to give the illusion of a dominant main axis. The growth form produced is called Pseudomonopodial .

True Monopodial Growth has its origin in Lateral Branching. Lateral Branches originate subapically and are associated with Leaves on stems.

Lateral roots have an internal origin near the Stele. The main root is called the Primary Root while the Branches are called Secondary Roots.

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Isotomous Branching in Marchantia

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Isotomous Branching (Lycopodium)

 

 

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Anisotomous Branching (Selaginella)


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Flax stems are relatively unbranched until they produce Flowers.
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One outcome of Monopodial growth!

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Lateral Branching in Roots produces a Monopodial System

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