WB01705.gif (1157 bytes)  Epidermis-4   WB01704_.gif (383 bytes)

The Epidermis of flowering plants produces a wide array ofYungLefTrich200.jpg (21945 bytes) hairs or Trichomes. They range from simple cells to elaborate,multi cellular, organ-like structures. Their adaptive significance is truly great, and many of their features are crucial for life on land. In some cases the relationship between plants and animals, as well as microbes is mediated by epidermal hairs. Young leaves are often covered with Trichomes. These may be ephemeral or persistent. They  provide protection for these primordial structures, but in most cases the true nature of their effects are unknown.
The most familiar, and most simple Trichomes are rootRootHairsWhole.jpg (19836 bytes) hairs! These are under-appreciated in terms of their functional significance. They are single cells! that greatly increase the absorptive surface area of roots. The most notorious interrelationship of root hairs and microbes occurs in  the legume-Rhizobium symbiosis. The root hairs are the initial recognition sites for the attachment of the nitrogen-fixing bacteria.

    

CottonFruJustOpen200.jpg (6160 bytes)
Dehiscent Cotton Fruit - The ovary wall has reflexed to reveal the tightly packed cotton fibers.
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Mature, Expanded Cotton Fruit showing the profusion of fibers.
The most important plant textile is cotton. Cotton fibers are gigantic cells. They are produced by the Epidermis of the Ovule. The ovule becomes the seed. The adaptive value of the fibers is probably due to enhanced seed dispersal. The fibers are pure cellulose and are ideal for the production of clothing.
CottonFibDiagLong400.jpg (6543 bytes) The strength of cotton fibers is due to the alternating layers of highly organized cellulose which have opposing pitches.
CottonFibXS200.jpg (6382 bytes) Cross-section of a cotton fiber showing the laminated nature of the cell wall.

The genus of cotton is Gossypium.

CottonHaw400.jpg (32178 bytes) Hawaiian cotton has been used to introduce valuable genes into commercial lines. This shows the potential genetic value of rare species.
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