Leaf
Anatomy - Lamina or Blade |
The Adaxial Epidermis
generally has a thick cuticle on the upper (adaxial) surface. Secretory &
Non-secretory Trichomes may be present. Stomata may occur on the upper epidermis but these
are most frequent on the lower (abaxial) surface. The Abaxial Epidermis usually has a
thin cuticle. However, it may have a dense layer of trichomes which increase the
"boundary layer" effect which decreases the rate of transpiration. Stomata are
routinely present on the abaxial surface and are usually more abundant than on the adaxial
surface. |
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Lamina from Ohia' lehua Metrosideros polymorpha showing a thick
adaxial cuticle (C) and abaxial Trichomes (T) |
Same but with Polarizing Optics. This brings out trichomes, crystals and
cells with thick, highly organized Cell Walls |
Mesophyll |
Palisade mesophyll is composed of columnar cells that can be one to several layers thick. It is most common
on the adaxial side of the leaf where sunlight is usually most abundant. Palisade
parenchyma are more efficient for photosynthesis and water retention because of their
large surface areas but small intercellular air spaces. |
SEM of Palisade Parenchyma
Palisade in a Sun Leaf
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Spongy mesophyll is usually found on the abaxial side of the leaf.
It may consist of stellate parenchyma or chains of elongate cells which have large,
lateral intercellular spaces. |
SEM of Spongy Mesophyll
Note the large air spaces! The cells have a stellate appearance and resemble the jacks used by children.
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Some plants have a uniform mesophyll which is composed of relatively isodiametric
cells. |
Mesophyll Cells in Wheat do not have pronounced shapes,
and give a uniform appearance. |
Nerium oleander (Oleander) has pronounced Palisade and
Spongy Layers |
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