Internal Leaf Anatomy

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General Features of the Lamina (Blade)

Three Tissue Systems are present!  

Epidermal - Ground (Mesophyll in leaves) - Vascular (Veins)

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Typical Dicot Leaf

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Cross Section of Sugar Cane Leaf (Monocot)

Leaf Anatomy - Lamina or Blade

The Adaxial Epidermis generally has a thick cuticle on the upper (adaxial) surface. Secretory & Non-secretory Trichomes may be present. Stomata may occur on the upper epidermis but these are most frequent on the lower (abaxial) surface.

The Abaxial Epidermis usually has a thin cuticle. However, it may have a dense layer of trichomes which increase the "boundary layer" effect which decreases the rate of transpiration. Stomata are routinely present on the abaxial surface and are usually more abundant than on the adaxial surface.

OhiaPubPol.jpg (47636 bytes)

Lamina from Ohia' lehua Metrosideros polymorpha showing a thick adaxial cuticle (C) and abaxial Trichomes (T)

Same but with Polarizing Optics. This brings out trichomes, crystals and cells with thick, highly organized Cell Walls

Mesophyll

Palisade mesophyll is composed of columnar cells that can be one to several layers thick. It is most common on the adaxial side of the leaf where sunlight is usually most abundant. Palisade parenchyma are more efficient for photosynthesis and water retention because of their large surface areas but small intercellular air spaces.

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SEM of Palisade Parenchyma

 

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Palisade in a Sun Leaf

Spongy mesophyll is usually found on the abaxial side of the leaf. It may consist of stellate parenchyma or chains of elongate cells which have large, lateral intercellular spaces.

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SEM of Spongy Mesophyll
Note the large air spaces! The cells have a stellate appearance and resemble the jacks used by children.

Some plants have a uniform mesophyll which is composed of relatively isodiametric cells. WheatMesoMM300.jpg (25982 bytes)
Mesophyll Cells in Wheat  do not have pronounced shapes, and give a uniform appearance.

Nerium oleander (Oleander) has pronounced Palisade and Spongy Layers

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