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Lycophyta-Selaginellales- 4

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The Leaves are Microphylls and usually have one vein/leaf. A couple of species have branched venation, however.

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Cross Section of a Selaginella Leaf

The Mesophyll is usually simple but it ranges from Uniform Lacunate (uniform means isodiametric cells & lacunate indicates prominent air spaces) to Palisade & Spongy. The latter is the characteristic anatomy for dicots and is considered to be highly advanced as the Palisade & Spongy tissues perform different functions. Homogenous Mesophyll would be ancestral or "primitive".

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Cross Section of an Olive Leaf (Dicot)

Most species have typical chloroplasts, but one species has large cup-shaped chloroplasts. This could be an interesting research project for someone interested in plastid evolution.

Stomata are present. The lack of stomata on the stem indicates that the Leaves are the primary sites of Carbon Fixation. This is a further step in the specialization of the aerial system with regards to the primary locus of photosynthesis.

Selaginella has a Ligule.This is a distinctive trait. They are flask-like glandular structures that can be found at the base of each leaf.The appear to secrete mucilage which protects  young leaves from drying as they expand.

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Ligule

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