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These are very similar to plants in the Marchantiales. They are generally less complex in their vegetative anatomy than Marchantia and its immediate relatives.

They are not very diverse as there are approximately 20 genera and 500 species.

Habitat = Very Wet

Their Growth Form is simple & Thallose.
They  may be
Unbranched or have  Dichotomous Branching.

ThallusBranch240.jpg (22505 bytes)

Branches may also emerge from the Midrib. This is sometimes irregular in appearance.

Key Anatomical Features
Lack Air Chambers

MonocleaThallus.jpg (25582 bytes)
This thallus of Monoclea resembles a Green Alga (Chlorophyta)

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Pelliaepiphylla Thalli

 

Lack Scales
Gametangia = Sessile
The Seta elevates the Capsule above the Thallus

The Pelliaceae & Pallaviciniaceae are the two most important families in this Order

An Apical Cell constitutes the Apical Meristem.

Dichotomous Branching can occur at the Apex.
The Ventral Mid Rib can also produce Branches

Thallus

The Thallus is Smooth & Translucent.

Unlike Marchantia there is Little Tissue Differentiation.

All Cells are Chlorophyllous and they superficially resemble Chorophytes.

There is not the level of tissue differentiation seen with Marchantia.

A Thickened "Midrib" is present.

The Midrib contains Xylem-Like Tissue!

PallaviciniaThalus240.jpg (41817 bytes)
Rhizoids are produced on the ventral side of the Midrib

metzgeriaventsamrhizoidsLab.jpg (54956 bytes)

 

 

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Lecture Topics