Protection
of the Sporophyte is provided by the Involucre & Calyptra. 
Pellia with Sporophytes
Development can be arrested during harsh
periods. This has obvious adaptive value because spore release can be delayed until
conditions are advantageous.

Pellia piphylla Sporophytes
The MeioSpores
are Chlorophyllous. This is thought to be an Ancestral trait because the resulting Gametophyte will require
light in order to survive.

Podometrium Spores
There are some mitotic divisions before the spore germinates this is
called Endosporic Development & is considered
to be an advanced trait because a multicellular organism has
a greater chance to survive & mature than a unicellular one.
The Spores are Heavy, however, and they
require free H2O for Dispersal. This is a
disadvantage! Why???
Elaters are present and in some cases they
are Attached to inner Capsule Wall.
They can have an Egg Beater-like action which
helps to disperse the Spores.

Lecture Topics