At maturity the Megagametophyte wall fractures along the Triradiate Ridge.
This exposes the Necks of
the Archegonia.
Fertilization
requires the presence of free water so that the Sperm can swim to the Egg.
Embryo
development is Endoscopic.
It grows into
the Megagametophyte and derives nutrition from the Storage Lipids. It is also protected from desiccation and other injuries by the thick Spore Wall.
The Shoot Apex eventually grows out of the
opening in the Megaspore Wall.
A
Root Apical Meristem forms at the pole opposite the Shoot Apical Meristem.
The Foot remains like an haustorium in the food
reserve and the Megagametophyte remains attached to the young Sporophyte for some time.
The
Overall pattern of Development resembles that of a Seed.
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![MegagametophyteEmb-1.jpg (51497 bytes)](MegagametophyteEmb-1_small.jpg)
Embryo Development is Endoscopic
![EmbEndo-2.jpg (47297 bytes)](EmbEndo-2_small.jpg)
The Shoot apex eventually responds to gravity and grows out of the Megagametophyte.
![GametoSporo240Lab.jpg (33238 bytes)](GametoSporo240Lab_small.jpg)
![Youngsporophyte.jpg (54083 bytes)](Youngsporophyte_small.jpg)
Sporophyte with discernable Root & Shoot: still attached to the Megagametophyte
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