The Gametophyte is small,
cylindrical & subterranean. It has an Apical Cell and Dichotomous Branching. It has Rhizoids & it has Gemmae
that are similar to those on the Sporophyte
Rhizome.
It also has Mycorrhizal
Fungal associations which are vital for its survival because it is not
photosynthetic. It has Vascular Tissue but this is
poorly and unevenly developed. Due to its small size and the fact that most of its cells
are pervaded by fungal hyphae, a Vascular System may be redundant. The Vascular System is,
however, a Protostele just like the Rhizome.
Antheridia are Superficial
and appear as small globes on the Gametophyte.
They produce multicellular Sperm which are Fern-like.
The Necks of the Archegonia
protrude above the surface of the Gametophyte but the Eggs are
embedded in the Gametophyte Tissues.