An updated classification of the Monocotyledoneae

Robert F. Thorne

1500 N. College Ave., Rancho Santa Ana Botanic Garden
Claremont, CA 91711, U.S.A.

with nomenclatural additions by

James L. Reveal

Norton-Brown Herbarum, Rm. 1211 H.J. Patterson Hall
University of Maryland
College Park, MD 20742-5815, U.S.A.



Since publication in 1992 by Botanical Review of my classification of the Angiospermae [Thorne, R.F. 1992b. Classification and geography of the flowering plant. Bot. Rev. 58: 225-348], many new contributions to their understanding have been made, especially for the Monocotyledoneae. This updated classification of that group is based on those contributions, especially those presented at the International Symposium "Monocotyledons: Systematics and Evolution" held at the Royal Botanic Garden, Kew, England in July 1993 and published in two volumes under the same title by the Royal Botanic Garden, Kew (Rudall et al. 1995). This updated classification is dedicated to my good friend Rolf Dahlgren.

The biggest changes in my monocotyledon classification since 1992 are based upon the highly informative results produced by the molecular taxonomists through analyses of DNA sequence data from the chloroplast gene rbcL, other plastid genes, chloroplast restriction analysis of the inverted repeat, and nuclear rRNA sequencing, combined with morphological and other data. One caveat here is that most molecular trees based on sequence data, no matter how parsimonious, are single character trees. Morphology in all its aspects from micromorphology to embryology, palynology, seed, seedling, fruit, floral, leaf, and stem anatomy remain hugely important, especially with proper weighting. Some of the most important features are the ultraviolet fluorescence behavior of cell walls, sieve-element plastids, and epicuticular wax crystalloids, pollen sulcus type, ovule structure, and seed coat and seed storage tissues. Other valuable approaches are use of hostplants and their predators or parasites, plant geography, and comparative chemistry.

In the following synopsis of the Monocotyledoneae, the hierarchy consists in descending order of superorders (-anae), orders (-ales), suborders (-inae), families (-aceae), and subfamilies (-oideae). The names of all hierarchial ranks are based on the principle of priority. The numerals in parentheses as (7/72) indicates 7 genera, 72 species. In summary, I recognize in the Monocotyledoneae nine superorders, 24 orders, 16 suborders, 114 families, 82 subfamilies, 2778 genera, and 57,755 species.

[Note added by Reveal: I have added to Dr. Thorne's listing all of the authorships for suprageneric names as known to me currently. Names without authorships have yet to be checked for their validity. I have added full synonymy except at the rank of subfamily. Dr. Thorne's outline does not as yet lend itself to ascertaining where he would assign such names. Finally, I have numbered each entry.]

1. Triuridanae Thorne ex Reveal, 1992 (10/50)
    1. Triuridales Hook.f., 1873 (10/50)
            1. Petrosaviaceae Hutch., nom. cons., 1934 (1/3)
            2. Triuridaceae Gardner, nom. cons., 1934
                            Lacandoniaceae E. Martínes & Ramos, 1989
2. Acoranae Reveal, 1997 (10/71)
     1. Nartheciales Reveal & Zomlefer, 1998 (9/69)
                            Tofieldiales Reveal & Zomlefer, 1998
             1. Tofieldiaceae Takht., 1995 (5/28)
                            Japonoliriaceae Takht., 1996
             2. Nartheciaceae Fr. ex Bjurzon, 1846 (4/41)
                            Heloniadaceae J. Agardh, 1858
                            Lophiolaceae Nakai, 1943
     2. Acorales Reveal, 1996 (1/2)
             1. Acoraceae Martynov, 1820 (1/2)
3. Aranae Thorne ex Reveal, 1992 (109/ 3075)
    1. Arales Dumort., 1829 (109/3075)
            1. Araceae Juss., 1789, nom. cons. (105/3040)
                            Arisaraceae Raf., 1838
                            Caladiaceae Salisb., 1866
                            Callaceae Rchb. ex Bartl., 1830
                            Cryptocorynaceae J. Agardh, 1858
                            Dracontiaceae Salisb., 1866
                            Lentiscaceae Horan., 1843
                            Orontiaceae Bartl., 1830
                            Pistiaceae Rich. ex C. Agardh, 1822
                            Pothaceae Raf., 1838
                   1a. Gymnostachyoideae (1/2)
                   1b. Orontioideae Müll. Berol., 1860 (3/6)
                   1c. Pothoideae Engl., 1876 (4/880)
                   1d. Monsteroideae (Schott) Engl., 1876 (12/370)
                   1e. Lasioideae Engl., 1876 (10/55)
                   1f. Calloideae Endl., 1837 (1/1)
                   1g. Aroideae Arn., 1832 (73/1690)
            2. Lemnaceae Gray, 1821, nom. cons. (4/37)
                            Wolffiaceae Bubani, 1901-1902
                    2a. Lemnoideae Bab., 1843 (2/16)
                    2b. Wolffioideae Engl., 1892 (2/21)
4. Alismatanae Takht., 1967 (57/415)
                            Butomanae Takht. ex Reveal, 1992
    1. Alismatales Dumort., 1829 (34/200)
                            Butomales Hutch., 1934
                            Elodeales Nakai, 1950
                            Hydrocharitales Dumort., 1829
                            Vallisneriales Nakai, 1949
            1. Butomaceae Rich., 1815, nom. cons. (1/1)
            2. Limnocharitaceae Takht. ex Cronquist, 1981 (3/8)
             3. Alismataceae Vent., 1799, nom. cons. (12/80)
                            Damasoniaceae Nakai, 1943
            4. Hydrocharitaceae Juss., 1789, nom. cons. (18/110)
                            Blyxaceae (Aschers. & Gürcke) Nakai, 1949
                            Elodeaceae Dumort., 1829
                            Enhalaceae Nakai, 1943
                            Halophilaceae J. Agardh, 1858
                            Hydrillaceae Prantl, 1880
                            Najadaceae Juss., nom. cons., 1789
                            Stratiotaceae Link, 1829
                            Thalassiaceae Nakai, 1943
                            Vallisneriaceae Link, 1829
                   4a. Hydrocharitoideae Eaton, 1836 (16/105)
                   4b. Thalassioideae Aschers. & Gürcke, 1889 (1/2)
                   4c. Halophiloideae Aschers. & Gürcke, 1889 (1/4)
    2. Potamogetonales Dumort., 1829 (23/215)
                            Aponogetonales Hutch., 1934
                            Cymodoceales Nakai, 1943
                            Juncaginales Hutch., 1934
                            Posidoniales Nakai, 1943
                             Ruppiales Nakai, 1950
                            Scheuchzeriales B. Boivin, 1956
                            Zosterales Nakai, 1943
        1. Aponogetonineae (1/47)
            1. Aponogetonaceae J. Agardh, 1858, nom. cons. (1/47)
        2. Potamogetonineae Engl., 1898 (22/170)
            1. Scheuchzeriaceae F. Rudolphi, 1830, nom. cons. (1/1)
            2. Posidoniaceae Hutch., 1934, nom. cons. (1/9)
            3. Cymodoceaceae N. Taylor, 1909, nom. cons. (5/16)
            4. Ruppiaceae Horan. ex Hutch., 1934, nom. cons. (1/8)
            5. Juncaginaceae Rich., 1808, nom. cons. (4/15)
                            Heterostylaceae Hutch., 1934
                            Lilaeaceae Dumort., 1829
                            Maundiaceae Nakai, 1943
                            Triglochinaceae Chevall., 1827
                   5a. Juncaginoideae (Rich.) Arn., 1832 (3/14)
                   5b. Lilaeoideae (1/1)
            6. Potamogetonaceae Dumort., 1829, nom. cons. (3/90)
                            Hydrogetonaceae Link, 1829
            7. Zosteraceae Dumort., 1829, nom. cons. (3/19)
            8. Zannichelliaceae Dumort., 1829, nom. cons. (4/12)
5. Lilianae Takht., 1967 (1312/27300)
     1. Liliales Perleb, 1826 (65/1545)
                            Alstroemeriales Hutch., 1934
                            Colchicales Dumort., 1829
                            Melanthiales R. Dahlgren ex Reveal, 1992
                            Paridales Dumort., 1829
                            Smilacales Lindl., 1833
                            Trilliales Takht., 1997
                            Veratrales Dumort., 1829
            1. Campynemataceae Dumort., 1829 (2/4)
            2. Melanthiaceae Batsch, 1802, nom. cons. (12/97)
                            Chionographidaceae (Nakai) Takht., 1996
                            Heloniadaceae J. Agardh, 1858
                            Veratraceae Salisb., 1807
                            Xerophyllaceae Takht., 1996
            3. Trilliaceae Lindl., 1846, nom. cons. (2/70)
                            Paridaceae Dumort., 1827
            4. Alstroemeriaceae Dumort., 1829, nom. cons. (5/165)
            5. Colchicaceae DC., 1805, nom. cons. (21/240)
                            Bulbocodiaceae Salisb., 1866
                            Burchardiaceae Takht., 1996
                            Merenderaceae Mirb., 1804
                            Petermanniaceae Hutch., nom. cons.
                            Uvulariaceae A. Gray ex Kunth, 1843, nom. cons.
                   5a. Colchicoideae Burmeist., 1837 (15/220)
                   5b. Uvularioideae A. Gray, 1848 (5/19)
            6. Rhipogonaceae Conran & Clifford, 1985 (1/6)
            7. Smilacaceae Vent., 1799, nom. cons. (2/312)
            8. Philesiaceae Dumort., 1829, nom. cons. (2/2)
            9. Luzuriagaceae Lotsy, 1911 (2/5)
            10. Calochortaceae Dumort., 1829 (1/65)
            11. Liliaceae Juss., 1789, nom. cons. (11/545)
                            Erythroniaceae Martynov, 1820
                            Fritillariaceae Salisb., 1866
                            Liriaceae Batsch, 1786
                            Medeolaceae (S. Wats.) Takht., 1987
                            Tulipaceae Batsch, 1786
                   11a. Medeoloideae (2/6)
                   11b. Lilioideae Eaton, 1836 (9/540)
            12. Tricyrtidaceae Takht., 1997 nom. cons. prop. (4/33)
                            Compsoaceae Horan., 1834
                            Scoliopaceae Takht., 1996
    2. Taccales Dumort., 1829 (11/645)
                            Dioscoreales Hook.f., 1873
                            Tamales Dumort., 1829
            1. Dioscoreaceae R. Br., 1810, nom. cons. (3/600)
                            Tamaceae Martynov, 1820
                            Tamnaceae J. Kickx f., 1826
            2. Stenomeridaceae J. Agardh, 1858 (1/2)
            3. Trichopodaceae Hutch., nom. cons. (2/2)
                            Avetraceae Takht., 1997
            4. Pentastemonaceae Duyfjes, 1992 (1/2)
            5. Stemonaceae Engl., 1887, nom. cons. (3/30)
                            Croomiaceae Nakai, 1937
                            Roxburghiaceae Wall., 1832
            6. Taccaceae Dumort., 1829, nom. cons. (1/11)
    3. Burmanniales Heintze, 1927 (15/165)
            1. Burmanniaceae Blume, nom. cons. (9/95)
            2. Thismiaceae J. Agardh, 1858, nom. cons. (4/40)
            3. Corsiaceae Becc., 1878, nom. cons. (2/28)
     4. Orchidales Raf., 1815 (1221/24,945)
                            Agavales Hutch., 1934
                            Alliales Traub, 1972
                            Asparagales Bromhead, 1838
                            Asteliales Dumort., 1829
                            Narcissales Dumort., 1829
                            Iridales Raf., 1815
                            Amaryllidales Bromhead, 1840
                            Ixiales Lindl., 1835
        1. Orchidineae Rchb., 1841 (804/18,670)
            1. Boryaceae (Baker) M.W. Chase, Rudall & Conran, 1997 (2/12)
            2. Hypoxidaceae R. Br., 1814, nom. cons. (9/120)
            3. Orchidaceae Juss., 1789, nom. cons. (788/18,500)
                            Apostasiaceae Lindl., 1833
                            Cypripediaceae Lindl., 1833
                            Limodoraceae Horan., 1847
                            Neottiaceae Horan., 1834
                            Neuwiediaceae (Burns-Bal. & V.A. Funk) R. Dahlgren ex Reveal Hoogland, 1991
                            Vanillaceae Lindl., 1835
                   3a. Apostasioideae Horan., 1847 (2/18)
                   3b. Vanilloideae (15/230)
                   3c. Cypripedioideae Kostel., 1831 (5/150-170)
                   3d. Spiranthoideae
                   3e. Orchidoideae Eaton, 1836
                   3f. Epidendroideae Kostel., 1831
            4. Lanariaceae H. Huber ex R. Dahlgren & A.E. van Wyk, 1988 (1/1)
            5. Asteliaceae Dumort., 1829 (3/35)
            6. Blandfordiaceae R. Dahlgren & Clifford, 1985 (1/4)
        2. Iridineae Engl., 1898 (79/1785)
            1. Tecophilaeaceae Leyb., 1862, nom. cons. (7/32)
            2. Doryanthaceae R. Dahlgren & Clifford, 1985 (1/2)
            3. Ixioliriaceae (Pax) Nakai, 1943 (1/3)
            4. Iridaceae Juss., 1789, nom. cons. (70/1750)
                            Crocaceae Vest, 1818
                            Galaxiaceae Raf., 1836
                            Geosiridaceae Jonker, 1939, nom. cons.
                            Gladiolaceae Raf., 1838
                            Isophysidaceae (Hutch.) F.A. Barkley, 1948
                            Ixiaceae Horan., 1834
                   4a. Isophysidoideae (1/1)
                   4b. Nivenioideae (6/85)
                   4c. Iridoideae Eaton, 1836 (36/720)
                   4d. Ixioideae Rchb. ex Klatt, 1866 (27/945)
        3. Asphodelineae (36/630)
            1. "Xeronemataceae" Thorne, ined. (1/2)
            2. Xanthorrhoeaceae Dumort., 1829, nom. cons. (1/30)
            3. Asphodelaceae Juss., 1789, nom. cons. (15/510)
                            Aloaceae Batsch, 1802
                   3a. Asphodeloideae Burnett, 1835 (8/270)
                   3b. Alooideae Link, 1829 (7/240)
            4. Hemerocallidaceae R. Br., 1810 (11/47)
                            Dianellaceae Salisb., 1866
                            Phormiaceae J. Agardh, 1858
                   4a. Hemerocallidoideae Kostel., 1831 (1/15)
                   4b. Phormioideae (10/32)
            5. Johnsoniaceae Lotsy, 1911 (8/40)
        4. Amaryllidineae (175/3315)
            1. Anthericaceae J. Agardh, 1858 (8/285)
            2. Laxmanniaceae Bubani, 1901-1902 (14/180)
                            Lomandraceae Lotsy, 1911
                   2a. Lomandroideae (5/65)
                   2b. Laxmannioideae (9/115)
            3. Alliaceae J. Agardh, nom. cons. (17/800)
                            Cepaceae Salisb., 1866
                            Gilliesiaceae Lindl., 1826
                            Milulaceae Traub, 1972
                            Tulbaghiaceae Salisb., 1866
                   3a. Allioideae Herb., 1837 (6/700)
                   3b. Tulbaghioideae (1/22)
                   3c. Gilliesioideae (Lindl.) Arn., 1832 (10/75)
            4. Agapanthaceae Voigt, 1850 (1/9)
            5. Amaryllidaceae J. St.-Hil., 1805, nom. cons. (66/730)
                            Brunsvigiaceae Horan., 1834
                            Crinaceae Vest, 1818
                            Cyrtanthaceae Salisb., 1866
                            Galanthaceae G. Mey., 1836
                            Gethyllidaceae Raf., 1838
                            Haemanthaceae Salisb., 1866
                            Leucojaceae Batsch, 1786
                            Narcissaceae Juss., 1789, nom. cons.
                            Oporanthaceae Salisb., 1866
                            Pancratiaceae Horan., 1847
                            Strumariaceae Salisb., 1866
                            Zephyranthaceae Salisb., 1866
            6. Hyacinthaceae Batsch, 1786 (40/900)
                            Eucomidaceae Salisb., 1866
                            Lachenaliaceae Salisb., 1866
                            Ornithogalaceae Salisb., 1866
                            Scillaceae Vest, 1818
            7. Themidaceae Salisb., 1866 (12/60)
            8. Aphyllanthaceae Burnett, 1835 (1/1)
            9. Behniaceae Conran, M.W. Chase & Rudall, 1997 (1/1)
            10. Anemarrhenaceae Conran, M.W. Chase & Rudall, 1997 (1/1)
            11. Herreriaceae Endl., 1841 (2/9)
            12. Agavaceae Endl., 1841, nom. cons. (10/315)
                            Yuccaceae J. Agardh, 1858
                   12a. Yuccoideae Kostel., 1831 (4/45)
                   12b. Agavoideae Herb., 1837 (6/270
            13. Hesperocallidaceae Traub, 1972 (2/ 25)
                             Hostaceae B. Mathew, 1988
        5. Asparagineae (29/675)
            1. Asparagaceae Juss., 1789, nom. cons. (2/ca 200)
            2. Dracaenaceae Salisb., 1866 (1/100)
                            Sansevieriaceae Nakai, 1936
            3. Nolinaceae Nakai, 1943 (4/50)
            4. Ruscaceae Sprengl. ex Hutch., 1934, nom. cons. (3/8)
            5. Convallariaceae Horan., 1834 (18/215)
                            Aspidistraceae Endl., 1841
                            Ophiopogonaceae Endl., 1841
                            Peliosanthaceae Salisb., 1866
                            Polygonataceae Salisb., 1866
                            Tupistraceae Schnizl., 1846
            6. Eriospermaceae Endl., 1841 (1/102)
6. Pandananae Thorne ex Reveal, 1992 (23/1325)
                            Cyclanthanae Thorne ex Reveal, 1996
    1. Pandanales Lindl., 1833 (3/875)
            1. Pandanaceae R. Br., 1810, nom. cons. (3/875)
                            Freycinetiaceae Brongn. ex Le Maout & Decne., 1868
                   1a. Pandanoideae Burnett, 1835 (2/700)
                   1b. Freycinetioideae Kurz, 1867 (1/175)
    2. Cyclanthales J.H. Schaffn., 1929 (11/200)
            1. Cyclanthaceae Dumort., 1829, nom. cons. (11/200)
                   1a. Carludovicoideae (10/200)
                   1b. Cyclanthoideae Burnett, 1835 (1/1)
    3. Velloziales R. Dahlgren ex Reveal, 1992 (9/250)
            1. Velloziaceae Endl., 1841, nom. cons. (8/250)
                            Barbaceniaceae Arn., 1832
                   1a. Vellozoideae (2/125)
                   1b. Xerophytoideae Herb., 1837 (6/125)
            2. Acanthochlamydaceae (Shou C. Chen) P.C. Kao, 1989 (1/1)
7. Arecanae Takht., 1967 (194/2360)
    1. Arecales Bromhead, 1840 (189/2350)
                            Cocosales Nakai, 1930
            1. Arecaceae Schultz Sch., 1832, nom. cons. (189/2350)
                            Acristaceae O.F. Cook, 1913
                            Borassaceae Schultz Sch., 1832
                            Calamaceae Kunth ex Perleb, 1838
                            Ceroxylaceae O.F. Cook, 1913
                            Chamaedoreaceae O.F. Cook, 1913
                            Cocosaceae Schultz Sch., 1832
                            Coryphaceae Schultz Sch., 1832
                            Geonomataceae O.F. Cook, 1913
                            Iriarteaceae O.F. Cook & Doyle, 1913
                            Lepidocaryaceae O.F. Cook, 1913
                            Malortieaceae O.F. Cook, 1913
                            Manicariaceae O.F. Cook, 1910
                            Nypaceae Brongn. ex Le Maout & Decne., 1868
                            Palmae Juss., 1789, nom. cons. et nom. alt.
                            Phoenicaceae Burnett, 1835
                            Phytelephaceae Mart. ex Perleb, 1838
                            Pseudophoenicaceae O.F. Cook, 1913
                            Sabalaceae Schultz Sch., 1832
                            Sagaceae Schultz Sch., 1832
                            Synechanthaceae O.F. Cook, 1913
                   1a. Coryphoideae Burnett, 1835 (40/400)
                   1b. Nypoideae Engl., 1892 (1/1)
                   1c. Calamoideae Beilschm., 1833 (22/670)
                   1d. Ceroxyloideae Drude, 1877 (10/155)
                   1e. Arecoideae Burnett, 1835 (113/1115)
                   1f. Phytelephoideae Drude, 1887 (3/8)
    2. Dasypogonales (4/8)
            1. Dasypogonaceae Dumort., 1829 (4/8)
                            Baxteriaceae Takht., 1996
                            Kingiaceae Endl., 1838
                   1a. Dasypogonoideae (2/6)
                   1b. Kingioideae Meisn., 1842 (2/2)
    3. Hanguanales R. Dahlgren ex Reveal, 1992 (1/2-5)
            1. Hanguanaceae Airy Shaw, 1964 (1/2-5)
8. Hydatellanae Takht. ex Reveal, 1992 (2/10)
    1. Hydatellales Cronquist, 1980 (2/10)
            1. Hydatellaceae U. Hamann, 1976, nom. cons. (2/10)
9. Commelinanae Takht., 1967 (1061/23,150)
                            Bromelianae R. Dahlgren ex Reveal, 1992
                            Juncanae Takht., 1967
                            Pontederianae Takht. ex Reveal, 1992
                            Typhanae Thorne ex Reveal, 1992
                            Zingiberanae Takht. ex Reveal, 1993
     1. Commelinales Dumort., 1829 (63/795)
                            Haemodorales Hutch., 1934
                            Pontederiales Hook.f., 1873
                            Philydrales Dumort., 1829
            1. Commelinaceae R. Br., 1810, nom. cons. (40/650)
                            Cartonemataceae Pichon, 1946, nom. cons.
                            Ephemeraceae Batsch, 1802
                            Tradescantiaceae Salisb., 1807
                   1a. Cartonematoideae (2/12)
                   1b. Commelinoideae Eaton, 1836 (38/640)
            2. Haemodoraceae R. Br., nom. cons. (13/100)
                            Conostylidaceae (Pax) Takht., 1987
                            Dilatridaceae M. Roem., 1840
                            Wachendorfiaceae Herb., 1837
                            Xiphidiaceae Dumort., 1829
                   2a. Haemodoroideae Arn., 1832 (8/34)
                   2b. Conostylidoideae Lindl., 1846 (5/65)
            3. Pontederiaceae Kunth, 1816, nom. cons. (6/38)
                            Heterantheraceae J. Agardh, 1858
                            Unisemataceae Raf., 1837
            4. Philydraceae Link, 1821, nom. cons. (4/5)
    2. Cannales Dumort., 1829 (89/2200)
                            Amomales Lindl., 1835
                            Musales Reveal, 1997
                            Zingiberales Griseb., 1854
        1. Musineae (2/40)
            1. Musaceae Juss., 1789, nom. cons. (2/40)
        2. Strelitzineae (3/7)
            1. Strelitziaceae (K. Schum.) Hutch., 1934 (3/7)
        3. Lowiineae (1/10)
            1. Lowiaceae Ridl., 1924, nom. cons. (1/10)
        4. Heliconiinae (1/200)
            1. Heliconiaceae (A. Rich.) Nakai, 1941 (1/200)
        5. Zingiberineae (50/1375)
            1. Zingiberaceae Lindl., 1835, nom. cons. (46/1275)
                            Alpiniaceae R. Br. ex F. Rudolphi, 1830
                            Amomaceae J. St.-Hil., 1805
            2. Costaceae Nakai, 1941 (4/100)
        6. Cannineae (32/570)
            1. Cannaceae Juss., 1789, nom. cons. (1/10-25)
             2. Marantaceae Petersen, 1888, nom. cons. (31/550)
    3. Bromeliales Dumort., 1829 (75/2820)
                            Mayacales Nakai, 1943
            1. Bromeliaceae Juss., nom. cons. (58/2730)
                            Tillandsiaceae Wilbr., 1834
                   1a. Brocchinioideae (1/21)
                   1b. Tillandsioideae Burnett, 1835 (10/1250)
                   1c. Bromelioideae Burnett, 1835 (32/725)
                   1d. Pitcairnioideae (15/735)
            2. Rapateaceae Dumort., 1829, nom. cons. (16/80)
                   2a. Rapateoideae Meisn., 1842 (7/24)
                   2b. Saxofridericioideae (9/55)
            3. Mayacaceae Kunth, 1842 (1/4-10)
    4. Xyridales Lindl., 1846 (14/1500)
                            Eriocaulales Nakai, 1930
        1. Xyridineae (5/325)
            1. Xyridaceae C. Agardh, 1823, nom. cons. (5/325)
                            Abolbodaceae Nakai, 1943
                   1a. Xyridoideae Kunth ex Arn., 1832 (2/300)
                   1b. Abolbodoideae (3/23)
        2. Eriocaulineae (9/1175)
            1. Eriocaulaceae P. Beauv. ex Desv., 1828 (9/1175)
    5. Juncales Dumort., 1829 (112/5295)
                            Cyperales Hutch., 1934
            1. Prioniaceae (1/1)
            2. Thurniaceae Engl., 1907, nom. cons. (1/3)
            3. Juncaceae Juss., 1789, nom. cons. (6/345)
            4. Cyperaceae Juss., 1789, nom. cons. (104/4945)
                            Caricaceae Burnett, 1835
                            Kobresiaceae Gilly, 1952
                            Papyraceae Burnett, 1835
                            Scirpaceae Batsch, 1786
                   4a. Mapanioideae (14/140
                   4b. Cyperoideae Kostel., 1831 (71/2345)
                   4c. Sclerioideae Burmeist., 1837 (15/335)
                   4d. Caricoideae Kostel., 1831 (5/2125)
    6. Typhales Dumort., 1829 (2/25)
            1. Typhaceae Juss., 1789, nom. cons. (2/25)
                            Sparganiaceae F. Rudolphi, 1830, nom. cons.
                   1a. Sparganioideae Link, 1829 (1/14)
                   1b. Typhoideae Link, 1829 (1/8-13)
    7. Restionales Hook.f., 1873 (719/10520)
                            Avenales Bromhead, 1838
                            Centrolepidales R. Dahlgr. ex Takht., 1997
                            Poales Small, 1903
        1. Restionineae (61/535)
            1. Anarthriaceae D.F. Cutler & Airy Shaw, 1965 (1/7)
            2. Restionaceae R. Br., 1810, nom. cons. (53/490)
                            Elegiaceae Raf., 1838
            3. Ecdeiocoleaceae D.F. Cutler & Airy Shaw, 1965 (2/2)
            4. "Hopkinsiaceae" (1/2)
            5. "Lyginiaceae" (1/1)
            6. Centrolepidaceae Endl., 1836 (3/35)
        2. Poineae (658/9980)
            1. Flagellariaceae Dumort., 1829, nom. cons. (1/4)
            2. Joinvilleaceae Toml. & A.C. Sm., 1970
            3. Poaceae (R. Br.) Barnh., 1895, nom. cons. (656/9975)
                            Aegilopaceae Martynov, 1820
                            Agrostidaceae Burnett, 1835
                            Alopecuraceae Martynov, 1820
                            Andropogonaceae Martynov, 1820
                            Anomochloaceae Nakai, 1943
                            Anthoxanthaceae (Link) Pfeiff., 1872
                            Arundinaceae (Dumort.) Hochst., 1850
                            Arundinellaceae (Stapf) Herter, 1940
                            Asperellaceae Link ex Pfeiff., 1872
                            Avenaceae Martynov, 1820
                            Bambusaceae Burnett, 1835
                            Chaeturaceae Link ex Pfeiff., 1872
                            Chloridaceae Herter, 1940
                            Coleanthaceae (Link) Pfeiff., 1873
                            Cynosuraceae (Link) Pfeiff., 1873
                            Echinariaceae (Link) Pfeiff., 1873
                            Ehrhartaceae (Link) Pfeiff., 1874
                            Eragrostidaceae (Stapf) Herter, 1940
                            Festucaceae Spreng. ex Pfeiff., 1873
                            Glyceriaceae (Link) Pfeiff., 1874
                            Gramineae Juss., 1789
                            Hordeaceae Burnett, 1835
                            Lepturaceae (Dumort.) Herter, 1940
                            Melicaceae Martynov, 1820
                            Miliaceae Burnett, 1835
                            Nardaceae Martynov, 1820
                            Oryzaceae Burnett, 1835
                            Panicaceae Voigt, 1845
                            Pappophoraceae (Kunth) Herter, 1940
                            Parianaceae Nakai, 1943
                            Paspalaceae (Link) Pfeiff., 1873
                            Phalaridaceae Burnett, 1835
                            Pharaceae (Stapf) Herter, 1940
                            Saccharaceae Martynov, 1820
                            Spartinaceae Burnett, 1835
                            Sporobolaceae Herter, 1941
                            Stipaceae Burnett, 1835
                            Streptochaetaceae Nakai, 1943
                            Triticaceae Hochst., 1848
                            Zeaceae A. Kern., 1891
                   3a. Anomochlooideae Pilg ex Potztal, 1957 (2/4)
                   3b. Pharoideae (Stapf) L.G. Clark & Judz., 1996 (3/12)
                   3c. Bambusoideae Luerss., 1893] (65/965)
                   3d. Ehrhartoideae Link, 1827 (14-18/150)
                   3e. Pooideae Benth., 1861 (154/3275)
                   3f. Arundinoideae Burmeist., 1837 (49/605)
                   3g. Danthonioideae (19/275)
                   3h. Aristidoideae (1/250)
                   3i. Chloridoideae Burmeist., 1837 (130/1125)
                   3j. Centothecoideae (10/25)
                   3k. Panicoideae Link, 1827 (207/3290)

Posted: 29 Mar 1999
Maintained by James L. Reveal
Norton-Brown Herbarium (MARY)
E-mail: jr19@umail.umd.edu